M. L.  Botton, Jr, C. N. Shuster, Sekiguchi, K., and Sugita, H., ?Amplexus and mating behavior in the Japanese horseshoe crab, Tachypleus tridentatus?, Zoological ScienceZoological science, vol. 13, pp. 151-159, 1996.

L. I.  Anderson and Jr, C. N. Shuster, ?Throughout geologic time: where have they lived?, The American Horseshoe Crab. Harvard University Press, CambridgeThe American Horseshoe Crab. Harvard University Press, Cambridge, pp. 189-223, 2003.